Tuesday, 14 June 2011

AUTONOMIC NERVOUS SYSTEM(ANS)

ANATOMY OF THE ANS

The ANS is  the major involuntary portion of  the nervous system and is  responsible for auto-
matic, unconscious bodily functions, such as control of heart rate and bloo  d pressure and both
gastrointestinal and genitourinary functions. The ANS is divided into two major subcategories:
the parasympathetic autonomic nervous system (PANS) and the sympathetic autonomic ner-
vous system (SANS).


Location of ANS Ganglia

Both the PANS and SANS have relay stations, or ganglia, between the CNS and the end organ,
but  the  somatic system does not. An  important  anatomic  difference between  the SANS and
PANS  is that the ganglia of the former  lie in two paraventral chains adjacent to the vertebral col-
umn, whereas most of  the ganglia of  the PANS system are located  in  the organs innervated.
 the major features of the ANS and the somatic systems and also shows the location of the major receptor types. These are:

NN-Nicotinic  receptors are located on cell bodies in ganglia of both PANS and SANS and in
the adrenal medulla.

NM-Nicotinic  receptors are located  on  the skeletal muscle motor end  plate  innervated  by
somatic motor nerves.

M1~3-Muscarinic  receptors are located on all organs and tissues innervated by postganglionic
nerves of the PANS and on thermoregulatory sweat glands innervated by the SANS.


Neurotransmitters

Acetylcholine (ACh) is the neurotransmitter at both nicotinic and muscarinic receptors  in tis-
sues  that are  innervated. Note  that all  direct transmission  from  the CNS  (preganglionic and
motor) uses ACh, but postganglionic transmission  in  the SANS  system may use one of the
organ-specific transmitters described below.

Norepinephrine  (NE) is  the  neurotransmitter  at most adrenoceptors in organs, as well as  in
cardiac and smooth muscle.

Dopamine  (DA) activates Dl  receptors, causing vasodilation in  renal and mesenteric vascular
beds.

Epinephrine  (E, from adrenal medulla) activates most adrenoceptors and is transported in the
blood.

BLOOD PRESSURE CONTROL MECHANISMS
 

Autonomic Feedback Loop

Blood pressure  is  the product of  total peripheral  resistance  (TPR) and cardiac  output (CO).
Both branches of the ANS are involved in the autonon~ic  (or neural) control of blood pressure
via feedback mechanisms. Changes  in mean  blood pressure  are detected  by  baroreceptors,
which  relay information  to the cardiovascular  centers in  the brainstem controlling PANS and
SANS outflow. For example, an increase in mean blood pressure elicits baroreceptor discharge,
resulting in increased PANS activity,  leading to bradycardia and decreased SANS activity, which
leads,  in turn, to decreased heart rate, force of contraction, and vasoconstriction. The resulting
decreases in cardiac output and total peripheral  resistance contribute  to  restoration of  mean
blood pressure toward its normal level. Conversely, decreases in blood pressure elicit ANS neu-
ral feedback involving decreased PANS outflow and increased SANS activity-actions leading
to increases  in cardiac output and total peripheral resistance.







Hormonal Feedback Loop

Blood pressure is also regulated via  the horrnonal  feedback loop shown in Figure 11-1-3. The
system  is  affected  only by  decreases in mean  blood pressure  (hypotension), which result  in
decreased renal blood  flow. Decreased renal pressure causes the  release of  renin, which pro-
motes  formation  of  the angiotensins. Angiotensin  I1  increases  aldosterone  release  from  the
adrenal cortex, which, via its mineralocorticoid actions to retain sodium  and water, increases
blood volume. Increased venous return results in an increase in cardiac output. Angiotensin  I1
also causes vasoconstriction, resulting in an increase in TPR,


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